University of Isfahan Taxonomy and Biosystematics 3115-9001 13 48 2021 09 23 48 شناسنامه علمی شماره 48 26481 10.22108/tbj.2021.26481 EN Journal Article 2022 03 19 https://tbj.ui.ac.ir/article_26481_0fa368715c1761bd7c84ae2af641179c.pdf